Table 1. Average percentage of foraging bouts (mean ± s.e., *n* = 8 bees) in which bees followed the optimal travel distance or linked flower visits by making nearest-neighbour movements in relation to the first visited flower. Numbers in parentheses indicate the travel distance for each sequence. Bees did not follow an optimal route if their first visit was not to either flower 1 or 6. Wilcoxon tests (*p*-values) were used to compare the percentage of bouts starting at each flower in which bees followed an optimal route or the nearest-neighbour route.