Table 1.

Average percentage of foraging bouts (mean ± s.e., n = 8 bees) in which bees followed the optimal travel distance or linked flower visits by making nearest-neighbour movements in relation to the first visited flower. Numbers in parentheses indicate the travel distance for each sequence. Bees did not follow an optimal route if their first visit was not to either flower 1 or 6. Wilcoxon tests (p-values) were used to compare the percentage of bouts starting at each flower in which bees followed an optimal route or the nearest-neighbour route.

first visited floweroptimizing overall travel distancelinking nearest-neighbour flowers
sequence% of boutssequence% of boutsp-value
1123 456 (2462 cm)11.41 ± 1.41124 563 (3380 cm)0.31 ± 0.060.02
2213 456 (2692 cm)1.41 ± 0.25214 563 (3518 cm)00.17
3345 621 (2889 cm)0.16 ± 0.05345 126 (2984 cm)1.56 ± 0.340.27
4453 216 (2855 cm)0.16 ± 0.05451 263 (3771 cm)0.47 ± 0.070.42
5543 216 (2633 cm)1.56 ± 0.29543 216 (2633 cm)1.56 ± 0.291
6654 321 (2462 cm)10 ± 0.90612 453 (2985 cm)0.16 ± 0.050.01