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This is an excellent study on the biomechanics of swimming in turtles. However, the most recent phylogenetic and paleontological analyses date the divergence of North American box turtles (Emydidae: Emydinae: /Terrapene/) to 14.5-11.5 Ma ([1-2]). To clarify, some extant Terrapene lineages likely diverged over the last five millions years [1-2]. Further, terrestrially or some degree thereof might be an ancestral feature of emydine turtles . Therefore, the assertion that terrestrially is recently evolved in emydids is debatable. Nonetheless, these observations would not alter the conclusion of functional convergence in tortoises and box turtles.
 Joyce, W.G., Petričević, A., Lyson, T.R. & Czaplewski, N.J. 2012 A new box turtle from the Miocene/Pliocene boundary (latest Hemphillian) of Oklahoma and a refined chronology of box turtle diversification. Journal of Paleontology 86, 177-190. (doi:10.1666/11-073.1).
 Spinks, P.Q., Thomson, R.C., McCartney-Melstad, E. & Shaffer, H.B. 2016 Phylogeny and temporal diversification of the New World pond turtles (Emydidae). Molecular Phylogenetics and Evolution 103, 85-97. (doi:10.1016/j.ympev.2016.07.007).
 Stephens, P.R. & Wiens, J.J. 2003 Ecological diversification and phylogeny of emydid turtles. Biol. J. Linn. Soc. 79, 577-610.
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