- Received November 12, 2015.
- Accepted January 20, 2016.
- Published online February 17, 2016.
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- 1School of Biological and Chemical Sciences, Queen Mary University of London, London, UK
- 2Raymond M. Alf Museum of Paleontology, Claremont, CA, USA
- 3Department of Anatomy, College of Osteopathic Medicine of the Pacific and College of Podiatric Medicine, Western University of Health Sciences, Pomona, CA, USA