- Received August 23, 2016.
- Accepted October 21, 2016.
- Published online November 16, 2016.
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© 2016 The Authors. Published by the Royal Society under the terms of the Creative Commons Attribution License http://creativecommons.org/licenses/by/4.0/, which permits unrestricted use, provided the original author and source are credited.
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- 1Messerli Research Institute, University of Veterinary Medicine Vienna, Medical University of Vienna, University of Vienna, Veterinärplatz 1, 1210 Vienna, Austria
- 2Department of Cognitive Biology, University of Vienna, Althanstr 14, 1090 Vienna, Austria
- 3Department of Zoology, University of Oxford, OX1 3PS Oxford, UK