- Received July 20, 2016.
- Accepted October 14, 2016.
- Published online November 9, 2016.
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© 2016 The Authors. Published by the Royal Society under the terms of the Creative Commons Attribution License http://creativecommons.org/licenses/by/4.0/, which permits unrestricted use, provided the original author and source are credited.
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- 1Department of Biological Sciences, Macquarie University, New South Wales 2109, Australia
- 2Department of Geology, University of California Davis, One Shields Avenue, Davis, CA 94568, USA
- 3Department of Geology and Geological Engineering, South Dakota School of Mines and Technology, Rapid City, SD 57701, USA
- 4Department of Earth Sciences, University of Oxford, South Parks Road, Oxford OX1 3AN, UK
- 5Department of Biology, University of York, Wentworth Way, York YO10 5DD, UK