- Received May 25, 2016.
- Accepted September 26, 2016.
- Published online October 25, 2016.
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© 2016 The Authors. Published by the Royal Society under the terms of the Creative Commons Attribution License http://creativecommons.org/licenses/by/4.0/, which permits unrestricted use, provided the original author and source are credited.
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- 1Ugyen Wangchuck Institute for Conservation and Environment Research, Bumthang, Bhutan
- 2Department of Migration and Immuno-Ecology, Max Planck Institute for Ornithology, 78315 Radolfzell, Germany
- 3Department of Biology, University of Konstanz, 87457 Konstanz, Germany
- 4Department of Civil, Environmental and Geodetic Engineering, Ohio State University, Columbus, OH 43210, USA